When sixteen-year-old Sally is evacuated to the English south coast, she is terrified by what lies ahead of her. All she knows are the sights and sounds of London's East End – but Sally swallows her tears as they leave the familiar landmarks behind, knowing that she has to be a Grown-Up Girl and play mother to her six-year-old brother Ernie. Playing mother is nothing new for Sally – their real mother Florrie, a good-time girl, hasn't even come to the station to wave them off and Ernie, crippled at an early age by polio, is used to depending on his older sister.

Cambridgeshire - March 1939. As the clouds of war begin to gather in Europe, the Talbot family of rural Blackberry Farm will be torn apart, just as so many families all over the world will be. Life will never be the same again. Whilst in London, the Salmons family will feel the pain of parting and loss. Brought together by war, the two families become intertwined and, as the outlook looks bleak, they must draw on each other’s strength to fight through the hard times.

The three sisters of Victory Walk in East London will face heartache and tears on the home front. As rationing, blackouts and bombs start to bite, life will change forever for the three sisters.

Alice Lake has arrived in London from Liverpool to start her training as a District Nurse, but her journey has been far from easy. Her parents think that she should settle down and get married, but she has already had her heart broken once and isn’t about to make the same mistake again. Alice and her best friend, Edith, are based in the East End, but before they’ve even got their smart new uniforms on, war breaks out, and Hitler’s bombs are raining down on London.

Daisy Wilder prefers the company of horses to people, bare feet and salt water to high heels and society parties. Then, in the dizzying aftermath of the attack on Pearl Harbor, Daisy enlists in a top secret program, replacing male soldiers in a war zone for the first time. Under fear of imminent invasion, the WARDs guide pilots into blacked-out airstrips and track unidentified planes across Pacific skies. But not everyone thinks the women are up to the job, and the new recruits must rise above and work side by side despite the resistance and heartache they meet along the way.

Three very different girls sign up as trainee nurses at a big London teaching hospital in 1934. Dora leaves her overcrowded, squalid working-class home for a better life. But has she got what it takes to keep up with other, better-educated girls? And will her hated stepfather ever let her go? Helen is born for the job; her brother is a doctor, her all-powerful mother a hospital trustee. But will Helen’s secret misery be her downfall? An aristocratic rebel, Millie’s carefree attitude will find her up in front of Matron again and again. Will she ever care enough to make a nurse? Or will she go back to the glamorous life she was born to?

1941. Audrey Coltrane has always wanted to fly. It's why she implored her father to teach her at the little airfield back home in Texas. It’s why she signed up to train military pilots in Hawaii when the war in Europe began. And it’s why she insists she is not interested in any dream-derailing romantic involvements, even with the disarming Lieutenant James Hart, who fast becomes a friend as treasured as the women with whom she flies. Then one fateful day, she gets caught in the air over Pearl Harbor just as the bombs begin to fall, and suddenly, nowhere feels safe.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.

December 1941: Evacuated from the threat of German bombs, sisters Amelie and Mireille have grown up under the storm clouds of war. Now they have joined the fight against Hitler, with Amelie training as a nurse to save wounded soldiers, and Mireille enlisted to help the Air Force wage war in the skies over Europe. But as each sister fights for peace, they are still haunted by the memory of their missing sister who was snatched away from them at the beginning of the war.

When Hazel is given the chance to parachute into Nazi-occupied France, she seizes the opportunity to do more for the British war effort than file paperwork. Alongside her childhood friend, French-born Rose, she quickly rises up the ranks of the freedom fighters. For Rose, the Resistance is a link to her late husband, and a way to move forward without him. What starts out as helping downed airmen becomes a bigger cause when they meet Sophia, a German escapee and fierce critic of Hitler who is wanted by the Gestapo.

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

Bristol 1936. Jenny Crawford has resigned herself to a loveless marriage living hand to mouth with their two children. Like many others, husband Roy struggles to find work at the docks but a chance encounter turns the family's fortunes around. Not only does he get regular work but they are also allocated a council house on Coronation Close on the outskirts of the city. Jenny and the children are overjoyed, this is the fresh start she could only ever dream of. But trouble feels never too far away.

Desperate to escape their stepfather’s house, sisters Amy and Lillian stow away aboard a train full of children being evacuated from London and the threat of Hitler’s bombs. Arriving in the seaside town of Worthing, they are taken in by kindly Norah and her husband, Jim. With their future now entrusted to strangers, can the girls finally find a safe harbour in these dark days of war? And will they find the strength to confront what they have been running from, when their past finally catches up with them?

The most ambitious of three sisters, Lorraine ‘Raine’ Linfoot always dreamed of becoming a pilot. As a spirited 17-year-old, she persuades her hero Doug Williams to teach her to fly. When war breaks out in 1939, Raine is determined to put her skills to good use. She enlists in the Air Transport Auxiliary, becoming one of a handful of brave female pilots flying fighter planes to the men on the front line.

When Sally, Beth, Margaret and Rachel meet at a job interview for the wonderful new store in Oxford Street, they have no idea they will become lifelong friends. When all four girls are lucky enough to be selected as sales staff their exciting new adventure begins. Join them as they overcome heartbreak and grief, find love and happiness and remain united in their friendship, whatever life throws at them.