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A winter sunrise over the great plains of Russia is no cause for celebration. The temperature barely rises above zero, and the guards at Penal Colony IK22 are determined to take their misery out on the prisoners—chief among them, one Zoya Zakharova. Once a master spy for Russian foreign intelligence, then the partner and lover of the Gray Man, she has information the Kremlin wants, and they don't care what they have to do to get it.

Like most spies, Jason Bourne lives in the shadows. But for Bourne, whose entire memory was stripped away by a bullet, those shadows hide dangerous secrets. When a desperate stranger in Zurich asks for his help, she calls him by the name David Webb, which is the identity he left behind years ago to become Jason Bourne. This woman tells him an incredible story–that her sister disappeared with David’s help after the deaths of four terrorists in the mountains. Now men from this same extremist group are trying to hunt her down.

US intelligence is reporting turmoil in the Russian navy. Their deadliest submarine, the Belgorod, has unexpectedly launched, and taken along with it a long list of questions. Who authorized the departure? What mission is it on? And, most disturbing of all, what weapons do the giant doors on the sub’s bow hide? It's been four decades since a similar incident with the Soviet sub, Red October, ended happily, thanks to a young CIA analyst named Jack Ryan.

From the glacial waters of Alaska to a sexy nightclub in the Bahamas, Treadstone agents are being hunted down and murdered. Someone high up in the U.S. government will stop at nothing to cover up a secret Treadstone mission from the past known as Defiance. With a team of killers hot on his trail, Jason Bourne chases the mystery of Defiance around the world. But as he closes in on the shocking truth, Bourne realizes that one man holds the key—his archenemy, the assassin known as Lennon.